JWSS - Journal of Water and Soil Science

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The variation and path coefficient analysis of bread-making quality traits have been studied for 145 genotypes of breeding lines, cultivars and landrace varieties of wheat (Triticum aestivum L.). Bread-making quality traits of genotypes was evaluated indirectly by protein percentage, SDS and Zeleny sedimentation tests, hardness index, test weight, bread volume, grain moisture, and water absorption percentage. Hardness index, Zeleny and SDS sedimentation tests had the highest coefficient of variability (13.51%, 11.83% and 11.03%, respectively). Factor analysis for the genotypes, based on qualitative traits revealed two factors, which explain 98.23% of total variation and were named grain protein index and bread volume factors. The result of correlation analysis indicated positive and significant relationship between protein percentage, SDS sedimentation test and other bread-making quality traits. On the first and second steps of stepwise regression analysis, protein percentage was the most effective trait in explaining different qualitative trait variations. Path analysis also showed the direct and significant effects of protein percentage, Zeleny sedimentation volume, grain moisture and water absorption percentage, and bread volume on SDS sedimentation test. According to cluster analysis based on qualitative traits, the cultivars and landrace varieties, from the view poit of protein quality and quantity, had the highest potential among other groups (generally breading lines).

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A diallel analysis of eight bread wheat (Ttriticum aestivum L.) cultivars was conducted to determine genetic parameters and the type of genetic control for yield and yield components. The parents and their F1 hybrids were evaluated in a randomized complete block design with 3 replications. General and specific combining ability effects were estimated by the method 2 of the Griffings model I, and the genetic parameters were estimated by the Jinks-Hayman method. According to the analysis of variance, the variances of parents and crosses were significant for all of the traits, except for the fertile tillers per plant in parents and harvest index in crosses. The variance of GCA was significant for almost all of the traits. GCA to SCA ratio indicated a large additive effects for all the traits, except for the number of fertile tillers, grain yield per plant and biological yield. Darab and Chamran cultivars were the excellent general combiners for days to heading. However, Falat for plant height, Arvand for number of fertile tillers, spike length and grain weight per spike, Qods and Arvand for spikeletes per spike, Falat and Arvand for grain yield per plant and Qods and Arvand cultivars for biological yield were the best general combiners. Based upon the Jinks-Hayman method, the average degree of dominance for grain weight per spike, 1000- grain weight, grain yield per plant and biological yield indicated that these traits might be controlled by over dominance effects and other traits by partial dominance. The correlation between Yr and (Wr + Vr) for the flag leaf length, spikeletes per spike, grains per spike and 1000-grain weight indicated that recessive allels enhanced these traits. But dominant allels had their contribution to enhance other traits. The narrow-sense heritability for grain yield per plant (4%) and biological yield (14%) and harvest index (37%) was lower. However, for other traits it was more than 50%. Therefore, it can be inferred that indirect selection of grain yield through selection for yield components such as spikeletes per spike, grains per spike and grain weight per spike with high heritability and correlation with grain yield can be more effective.

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Diallel analysis was used to estimate the combining ability, gene action, gene number, heritabilties and other genetic parameters of a set of wheat genotypes. For this purpose, nine parents and their 36 crosses were evaluated for 9 traits in a randomized complete block design with three replications in 1996. The analysis of variance revealed significant differences among all genotypes for all traits. Estimates of general and specific combining ability mean squares based on Griffing’s Method 2 indicated the importance of additive and non additive effects in the expression of all traits. Alvand and Roshan cultivars for grain yield per plant, Alvand for grain number per main spike and main spike weight, and Alvand and Alamoot for 1000 grain weight were the best combiners, thus use of them is beneficial for these traits. Based on the estimates of average degree of dominance and results of graphical analysis, the gene action for grain number and spikeletes per main spike were partial dominance, while for grain yield per plant, biological yield, plant height, harvest index and 1000 grain weight, overdominance gene actions were observed. Moreover, genotypic correlation coefficients of grain yield per plant with grain number per main spike, 1000 grain weight, grain weight per main spike and main spike weight were positive and significant.

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Inheritance of physiologicaly related salt tolerance traits including Na⁺ and K⁺ contents, K⁺/Na⁺ ratio of young leaves and biological yield (BY) in six basic generations (P₁, P₂, F₁, F₂, Bc₁ and Bc₂) and their reciprocal crosses derived from crosses between Kharchia × Niknejad and Shorawaki × Niknejad were studied in sand culture under high salinity treatment (EC = 22.5 dS m^-1). Generation means analysis indicated that a simple genetic model (including additive and dominance effects) is sufficient for Na⁺ and BY in Kharchia × Niknejad cross and for Na⁺, K⁺/Na⁺ ratio and BY in Shorawaki × Niknejad cross but, for other traits digenic interactions (additive × additive and dominance × dominance) were important parameters in the expression of salt tolerance of the various generations. Dominance genetic effects were predominant genetic components in most of the models. Weighted generation variances analysis suggested that dominance variance component was more important for Na⁺ and K⁺ content in both crosses. This result was confirmed by significant differences between back cross generations. Results also showed negligible dominance for K⁺/Na⁺ ratio in both crosses, multidirectional dominance for BY in Kharchia × Niknejad cross and absence of dominance gene action in Shorawaki × Niknejad. Kharchia × Niknejad cross showed, in general, more genetic variation, broad-and narrow sense heritabilities than Shorawaki × Niknejad cross, indicating the important role of primary differences between parents. These results revealed that recurrent selection followed by pedigree breeding or a selective diallel mating system may prove useful in improving salinity tolerance of wheat plants. The involvement of dominance × dominance interactions for some traits indicates that it is necessary to postpone selection for salt tolerance of wheat to advanced generations, when sufficient epistatic interactions have become fixed.